Professional Training and Experience

• DVM: Washington State University, College of Veterinary Medicine, 1971
• PhD: The Ohio State University, College of Veterinary Medicine, Department of Veterinary Pathobiology, 1974. Thesis title: "Immunologic Studies of Canine Distemper Virus Infection in Gnotobiotic Dogs."(Adalbert Koestner DVM, PhD, advisor)
• Diplomate: American College of Veterinary Pathology (ACVP) 1976.
• Professor, Department of Veterinary Biosciences, 1983 to present

Research Interests

• Pathogenesis of infectious diseases
• Canine and porcine immunology
• Comparative immunology, immunopathology, pathology and general pathology
• Gasteroenterology and mucosal immunology